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Which of the following bacterial cells (strains), acting as donors during conjugation, can play a role...

Which of the following bacterial cells (strains), acting as donors during conjugation, can play a role in transferring chromosomal DNA from the donor cell to the recipient cell?  

A) F+, hfr, F'

B) F+ and F-, only

C) Hfr and F', only

D) Hfr only

E) F' only

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