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Why might the response to selection (R) be less than the selection differential (S)?

Why might the response to selection (R) be less than the selection differential (S)?
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The selection differential is the difference of the base population mean and the mean of the selected parents. The selection response is how much gain you make when mating the selected parents.

Remember, the narrow sense heritability is a measure of the genetic component that is contributed by the additive genetic variance. The response to selection can thus be dervied by multiply the heriability by the selection differntial.

Selection differentials increase with angling intensity and the associated higher annual exploitation rates. Positive selection differentials would cause the mean phenotype to increase, whereas negative selection differentials would cause it to decrease.

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