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You measure the milk yield of a Holstein population. You know the variance of milk yield...

You measure the milk yield of a Holstein population. You know the variance of milk yield is 1000 lbs2 and the variance caused by permanent environmental effect is 230 lbs2. The correlation coefficient between milk yield and breeding value is 0.6. The repeatability of milk yield is 0.7. What is the variance caused by gene combination effect?
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